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Iirc the rods that see in grayscale but are good at light perception, have quicker response than cones that do color vision. However, rods are most engaged in low-light conditions, which here is true for the frog, so this part kinda contradicts the response time hypothesis.
I think the point is that there’s enough light around the center void circle to bypass that. I’m not that versed in this illusion tho.